Associate Professor at Leiden University


* Supervised M.Sc. or Ph.D. student paper

I = journal impact factor


Prochazkova, E., Sjak-Shie, E. E., Behrens, F., Lindh, D., & Kret, M. E.* (2019). The choreography of human attraction: physiological synchrony in a blind date setting. bioRxiv, 748707. Download.

Behrens, F., Snijdewint, J. A., Moulder, R. G., Prochazkova, E., Sjak-Shie, E. E., Boker, S. M., & Kret, M. E.* (2019). Physiological synchrony promotes cooperative success in real-life interactions. bioRxiv, 792416. Download.

Van Berlo, E., Bionda, T., & Kret, M.E.* (2020), Attention towards emotions is modulated by familiarity with the expressor. A comparison between bonobos and humans. bioRxiv. Download.

Behrens, F., & Kret, M. E.* (2020). Under the umbrella of prosocial behavior – a critical comparison of paradigms. PsyArxiv. Download.

Mattavelli, S., Brambilla, M., & Kret, M. E. It is written in the eyes: Inferences from pupil size and gaze orientation shape interpersonal liking. PsyArxiv. Download.


Van Berlo, E., Otten, M., Van de Ven, E., Binnekamp, J. & Kret, M.E.* (under review). A picture is worth a thousand words: Measuring implicit associations with a pictorial IAT.

Rotteveel, M., Kret, M.E., Winkielman, P., & van Dijk, W.W. (under review). On the parallel influences of facial and bodily expressions of emotion on social trust.

Van Berlo, E., Moczynski, A.C., Ploeger, A., & Kret, M.E.* (under review). Belongingness in great apes: exploring the relationship between social motivation and the emergence of theory of mind.

Danel, D.P., Wacewicz, S., Kleisner, K., Lewandowski, Z., Kret, M.E., Żywiczyński, P., & Perea-Garcia, J.O. (under review). Sex differences in ocular morphology in Caucasian people. Body Image.

In Press

Straffon, L., ea Visual attention bias for self-made artworks. (in press). Psychology of Aesthetics, Creativity, and the Arts.

Frisanco, A. All that Meets the Eye: The contribution of reward processing and pupil mimicry on pupillary reactions to facial trustworthiness. Current Psychology

Prochazkova, E., Sjak-shie, E.E., Behrens, F., & Lindh, D., & Kret, M.E.* Physiological synchrony predicts attraction in a blind date setting. Nature Human Behavior

Kret, M.E., & van Berlo. Attentional bias in humans towards human and bonobo expressions of emotion. Evolutionary Psychology, 19(3), 14747049211032816

Folz, J., Fiacchino, D., Nikolic, M., van Steenbergen, H., & Kret, M.E. (2021). Reading your emotions in my physiology? Reliable emotion interpretations in absence of a robust physiological resonance. Affective Science,

Aktar, E., Nimphy, C.A., Kret, M.E., Perez-Edgar, K.E., Bogels, S., & Raijmakers, M. (in press) Pupil responses to dynamic negative facial expressions of emotion in infants and parents” to Developmental Psychobiology. Developmental Psychobiology.

Aktar, E., Nimphy, C.A., Bogels, S., Raijmakers, M. Perez-Edgar, K.E., Kret, M.E. (in press). Attention Biases to Threat in Infants and Parents: Links to Parental and Infant Anxiety Dispositions. Research on Child and Adolescent Psychopathology. I = 2.290

Samara, I., Roth, T., & Kret, M.E.* (in press). The role of emotion projection, sexual desire, and self-rated attractiveness in the sexual overperception bias. archives of sexual behavior. I = 3.458.

Kret, M.E., Maitner, A.T., Fischer, A.H. (in press). Interpreting Veiled Emotions: A comparison between a Middle Eastern and Western-European sample. Journal of Nonverbal Behavior.

Roth, T., Samara, I., & Kret, M.E.* Multimodal mate choice: exploring the differential effects of sound, sight and scent on partner choice in a speed-date paradigm. Evolution and Human Behavior

Roth, T.S., Du, X., Samara, I., & Kret, M.E. (2021). Attractiveness modulates attention, but does not enhance gaze cueing. Evolutionary Behavioral Sciences, 10.1037/ebs0000265

Dukes, D., …Kret, M.E., …, Sander, D. (under review). Are we now in the era of affectivism? Nature Human Behaviour. # collaboration of numerous affective scientists. The order of authors is alphabetical except for the first and last.

Roth, T.S.*, Samara, I.*, Prochazkova, E., Tan, J., & Kret, M.E. (2021). A comparative framework of inter-individual coordination and pair-bonding. Current Opinion in Behavioral Sciences.

Wijnen, M., Massen, J., & Kret, M.E. (in press). Gender bias in the allocation of student grants. Scientometrics. I = 2.770.

Tsou, Y., Li, B., Kret, M.E., Wiefferink, C.H., Frijns, J.M., & Rieffe, C. (in press). Hearing status affects children’s emotion understanding in dynamic social situations: An eye-tracking study. Ear and Hearing. I =3.063

Nieuwburg, E., Ploeger, A., & Kret, M.E.* (2021). Emotion recognition in nonhuman primates: How experimental research can contribute to a better understanding of underlying mechanisms. Neuroscience & Biobehavioral Reviews. I = 8.330

Kim, Y., & Kret, M.E.* (in press). The emotional expressions and emotion perception in nonhuman primates. In Dukes, D., Samson, A.C., & Walle, E.A. (Eds.), The Oxford Handbook of Emotional Development. Oxford University Press.

Roth, T.S., Samara, I., & Kret, M.E.* (in press). Ultimate and proximate factors underlying sexual overperception bias: A reply to Lee et al. (2020). Evolution and Human Behavior. I = 2.959.


Dukes, D., et al. (2021). The rise of affectism. Nature Human Behavior, 5, 816-820.


Krippl, M., Kret, M.E., Dickson, J.M., Li, C.R., Lowe, L. (2020). A sensorimotor control framework for understanding emotional communication and regulation. Neuroscience and biobehavioral reviews, 112, 503-518.

Yamamoto, H.W., Kawahara, M. Kret, M.E., & Tanaka, A. (2020). Cultural Differences in Emoticon Perception: Japanese See the Eyes and Dutch the Mouth of Emoticons. Letters on Evolutionary Behavioral Science, 11(2), 40-45.. Download.

Behrens, F., Snijdewint, J. A., Moulder, R. G., Prochazkova, E., Sjak-Shie, E. E., Boker, S. M., & Kret, M. E. (2020). Physiological synchrony is associated with cooperative success in real-life interactions. Scientific reports10(1), 1-9. I = 3.998

van Berlo, E., Díaz-Loyo, A., Juárez-Mora, O., Kret, M.E., Massen, J. (2020). Experimental evidence for yawn contagion in orangutans (Pongo pygmaeus). Scientific Reports, 10(1), 1-11. I = 3.998

Tsou, Y., Li, B., Kret, M.E., Sabino da Costa, I., Rieffe, C. (2020). Reading emotional faces in deaf and hard-of-hearing and typically hearing children. Emotion. I = 3.039.

Massen, J. J., Bauer, L., Spurny, B., Bugnyar, T., & Kret, M. E. (2020). Reply to:“The data do not support the existence of an ‘Old Boy network’in science. Some critical comments on a study by Massen et al.”. Scientific Reports10(1), 1-3.

Laméris, D., van Berlo, B., Sterck, E., Bionda, T., & Kret, M.E.* (2020). Low relationship quality predicts scratch contagion during tense situations in orangutans (Pongo pygmaeus). The American Journal of Primatology, e23138. I = 2.009. Download.

Kret, M.E.*, & Roth, T. (2020). Anecdotes in animal behaviour. Behaviour (0)  1-2. Download.

Aktar, E., Raaijmakers, M., & Kret, M.E.* (in press). Pupil mimicry in infants and parents. Does parents’ empathy matter? Cognition and Emotion. I = 2.563.

Jaasma, L., Kamm, I., Ploeger, A. & Kret, M.E.* (2020). The exceptions that prove the rule? Spontaneous helping behaviour towards humans in some domestic dogs. Applied Animal Behaviour Science, 104941I = 1.817

Kret, M.E.*, Prochazkova, E., Sterck, E. H. M., Clay, Z. (in press). Emotional expressions in human and non-human great apes. Neuroscience Biobehavioral Reviews, 115, 378-395. I = 11.075.

Olvido Perea-García, J., Miani, A., Behrens, F., Kret, M.E., Mitkidis, P., & Kundtová Klocová, E. (In press). Low-level effects of perceived eye colour during eye contact. Physiology & Behavior.

Williams, J., et al. (in press). A sensorimotor control framework for understanding emotional communication and regulation. Neuroscience & Biobehavioral Reviews. I = 11.075.  Download.


Kret, M.E., Massen, J., & Bliss-Moreau, E. (2019). Special issue: Building a truly comparative affective science. Neuroscience and Biobehavioral Reviews. Download.

Ding, X.#, Liu, J.#, Kang, T., Wang, R., & Kret, M. E. (2019). Automatic change detection of emotional and neutral body expressions: Evidence from visual mismatch negativity. Frontiers in psychology10, 1909.

Kret, M.E., & de Dreu, C.K.W. (2019). The power of pupil size in establishing trust and reciprocity. JEP General. I = 4.420.

Behrens, F. & Kret, M.E.* (2019). The interplay between face-to-face contact and feedback on cooperation during real-life interactions. Journal of Nonverbal Behavior, 43(4), 513-528. I = 1.595. Download.

Olvido Perea-García, J., Kret, M.E., Monteiro, A., & Hobaiter, C. (2019). Scleral pigmentation leads to conspicuous, not cryptic, eye morphology in chimpanzees. Proceedings of the National Academy of Sciences,  116(39), 19248-19250. I = 10.4.

Brambilla, M., Biella, M., Kret, M.E. (2019). The power of pupils in predicting conforming behavior. Social Influence, 14(2), 40-49. I = 0.979.

Quesque, F., Behrens, F., Kret, M.E.* (2019). Pupils say more than a thousand words: Pupil size reflects how observed actions are interpreted. Cognition, 190, 93-98. I = 3.634. Download.

Kelsey, C., Krol, K., Kret, M.E., & Grossmann, T. (2019). Infants’ brain responses to pupillary changes in others are affected by race. Scientific Reports, 9 (1), 4317. I = 4.122.

Kret, M.E., & Sjak-Shie, E. (2019). Preprocessing pupil size data. Guideline and code. Behavior Research Methods, 1-7. I = 3.623. Download.


Brambilla, M., Biella, M., & Kret, M.E. (2018). Looking into your eyes: observed pupil size influences approach-avoidance responses. Cognition and Emotion, 33(3), 616-622. Download.

Prochazkova, E., Prochazkova, L., Giffin, M.R., Scholte, H.S., De Dreu, C.K.W., & Kret, M.E.* (2018). Reply to Mathôt and Naber: Neuroimaging shows that pupil mimicry is a social phenomenon. Proceedings of the National Academy of Sciences115(50), E11566-E11567. I = 10.4.

Prochazkova, E., Prochazkova, L., Giffin, M. R., Scholte, H.S., De Dreu, C.K.W., & Kret, M.E.* (2018). Pupil mimicry promotes trust through the theory-of-mind network. Proceedings of the National Academy of Sciences115(31), E7265-E7274. I = 10.4.

Koelkebeck, K., Vosseler, A., Kohl, W., Fasshauer, T., Lencer, R., Satoh, S., Kret, M.E., Minoshita, S. (2018). Masked ambiguity – emotion identification in schizophrenia and major depressive disorder. Psychiatry Research, 270, 852-860. I = 2.25

Kret, M.E., & Straffon, L. (2018). Reply to Crivelli et al.: The different faces of fear and threat. Evolutionary and cultural insights. Journal of Human Evolution, 125, 193-197. I = 3.767. Download.

Kret, M.E., Muramatsu, A., & Matsuzawa, T. (2018). Emotion processing across and within species. A comparison between humans and chimpanzees (Pan Troglodytes). Journal of Comparative Psychology, 132(4), 395. I = 2.242. Download.

Fischer, A., Broekens, J., & Kret, M.E. (2018). Gender differences in self-reported emotional intelligence: A test of the emotional sensitivity hypothesis. PlosOne, 13(1), e0190712. I = 3.730. Download.

Wehebrink, K.S., Koelkebeck, K., Piest, S., de Dreu, C.K.W., & Kret, M.E.* (2018). Pupil mimicry and trust – Implications for depression. Journal of Psychiatric Research, 97, 70-76. I = 2.528

Liedtke, C. Koelkebeck, K., & Kret, M.E.* (2018). Emotion recognition from faces with in- and out-group features in patients with depression. Journal of Affective Disorders, 227, 817-823. I = 3.845. Download.

Van Breen, J., de Dreu, C.K.W., & Kret, M.E.* (2018). Pupil to Pupil: effect of a partner’s pupil size on (dis)honest behavior. Journal of Experimental Social Psychology, 74, 231-245. I = 2.219. Download.

Kret, M.E., & Fischer, A. (2018). Recognition of facial expressions is moderated by Islamic cuesCognition and Emotion, 32(3) 623-631I = 2.688. Download.

Kret, M. E. (2018). The role of pupil size in communication. Is there room for learning? Cognition and Emotion, 32(5), 1139-1145. I = 2.688. Download.


Van Rooijen, R., Ploeger, A., & Kret, M.E.* (2017). The dot-probe task to measure emotional attention: A suitable measure in comparative studies? Psychonomic Bulletin & Review, 24(6), 1686-1717. I = 3.390. Download.

Massen, J. J., Bauer, L., Spurny, B., Bugnyar, T., & Kret, M. E. (2017). Sharing of science is most likely among male scientists. Scientific Reports7(1), 12927. I = 5.228. Download.

Prochazkova, E., & Kret, M.E.* (2017). Connecting minds and sharing emotions through mimicry: A Neurocognitive Model of Emotional Contagion. Neuroscience & Biobehavioral Reviews, 80, 99-114. I = 8.580. Download.

Kret, M.E., & de Dreu, C.K.W. (2017). Pupil-mimicry conditions trust in exchange partners: Moderation by oxytocin and group membership. The Royal Society B-Biological Sciences, 284(1850), 20162554. I = 5.051. Download.

Koelkebeck, K., Kohl, W., & Kret, M.E. (2017). Attachment style moderates theory of mind abilities in depression. Journal of Affective Disorders, 15(213), 156-160. I = 3.996. Download.

Koelkebeck, K., Uwatoko, T., Tanaka, J., & Kret, M.E. (2017). How culture shapes social cognition deficits in mental disorders- A review. Social Neuroscience, 12(2):102-112. I = 3.653. Download.

Kret, M.E., Roelofs, K., Stekelenburg, J.J., & de Gelder, B. (2017). From face to hand: attentional bias towards expressive hands in social anxiety. Biological Psychology, 122:42-50. I = 3.399. Download.

Kret, M.E. (2017). Spiegelen van pupil-grootte geeft vertrouwen. InMind Magazine: Psychologie voor iedereen. Download.

Kret, M. E., & Bocanegra, B. R., eds. (2017). Adaptive hot cognition: How emotion drives information processing and cognition steers affective processing. Lausanne: Frontiers Media. Download.


Kret, M.E., Jaasma, L., Bionda, T., & Wijnen, J.G.  (2016). Bonobos (Pan Paniscus) show an attentional bias towards conspecifics’ emotions. Proceedings of the National Academy of Sciences,113(14), 3761-3766. I = 9.674. Download.

Kret, M.E., & Tomonaga, M. (2016). Getting to the bottom of processing behinds in humans and chimpanzees (Pan Troglodytes). The face inversion effect revisited. PlosOne, 11(11): e0165357. I = 3.730. Download.

Kret, M. E., & Bocanegra, B. R. (2016). Editorial: Adaptive hot cognition: How emotion drives information processing and cognition steers affective processing. Frontiers in Psychology, 7:1920. I = 2.800. Download.

de Dreu, C.K.W., Kret, M.E., & Sauter, D. A. (2016). Assessing emotional vocalizations from cultural in-group and out-group depends on oxytocin. Social Psychological Personality Science, 7, 837-846I = 2.561. Download.

Stamkou, E., van Kleef, G., Fischer, A., & Kret, M.E. (2016). Are the powerful really blind to the feelings of others? How power and legitimacy shape emotion perception. Personality and Social Psychology Bulletin, 42(6), 755-768. I = 2.515. Download.

De Dreu, C.K.W., Kret, M.E., & Sligte, I. (2016). Modulating prefrontal control in humans reveals distinct pathways to competitive success and collective waste. Social Cognitive and Affective Neuroscience, 11(8):1236-1244. I = 7.372. Download.

De Dreu, C.K.W., & Kret, M.E. (2016). Oxytocin conditions intergroup relations through up-regulated in-group empathy, cooperation, conformity, and defense. Biological Psychiatry, S0006-3223(15)00259-0. I = 9.472. Download.


Kret, M.E. Fischer, A.H. & de Dreu, C.K.W. (2015). Pupil-mimicry correlates with trust in in-group partners with dilating pupils. Psychological Science, 26(9), 1401-1410. I = 4.43. Download.

Kret, M.E. & Ploeger, A. (2015). Emotion processing deficits: a liability spectrum providing insight into comorbidity of mental disorders. Neuroscience & Biobehavioral Reviews, 52, 153–171. I = 11.075. Download.

De Valk, J. M., Wijnen, J. & Kret,  M.E.* (2015). Anger fosters action. Fast responses in a motor task involving approach movements towards angry faces and bodies. Frontiers in Psychology, 6:1240.  I = 2.800. Download.

Kret, M.E. (2015). Emotional expressions beyond facial muscle actions. A call for studying autonomic signals and their impact on social perception. Frontiers in Psychology, 6, 711.  I = 2.800. Download.



Kret, M.E., Tomonaga, M. & Matsuzawa, T. (2014). Chimpanzees and humans mimic pupil-size of conspecifics. PlosOne, 9(8). I = 3.730. Download.

Kret, M.E.  Tomonaga, M. de Dreu, C.K.W., Fischer, A.H. & Matsuzawa, T. (2014). Role of pupil-synchronization in trust. International Journal of Psychophysiology, 94, 174.

Kret, M.E. (2014). Wanneer een lach een vuist wordt: De perceptie van gezicht- en lichaamsexpressies in gewelddadige delinquenten. De Psycholoog. Download.

Ten Velden, F.S. , Baas, M., Shalvi, S., Kret, M.E. &  De Dreu, C.K.W. (2014). Oxytocin mdifferentially modulates compromise and competitive approach but not withdrawal to antagonists from own versus rivaling other groups. Brain Research, 1580, 172-179. I = 2.957.  Download.


Kret, M.E., Roelofs, K., Stekelenburg, J.J., & de Gelder, B. (2013). Emotional signals from faces, bodies and scenes influence observers’ face expressions, fixations and pupil-size. Frontiers in Human Neuroscience, 7(810). I = 2.900. Download.

Kret, M. E. & de Dreu, C.K.W. (2013). Oxytocin-motivated ally selection is moderated by digit ratio and empathic concern. Frontiers in Neuroscience, 7(1). I = 2.900. Download.

Kret, M.E., Stekelenburg, J.J., Roelofs, K., & de Gelder, B. (2013). Perception of face and body expressions using electromyography, pupillometry and gaze measures. Frontiers in Psychology, 4(28). I = 2.800. Download.

Kret, M.E., & de Gelder, B. (2013). When a smile becomes a fist: The perception of facial and bodily expressions of emotion in violent offenders. Experimental Brain Research, 228(4):399-410. I = 2.296. Download.


Kret, M.E. (2012). De invloed van een niqab op het herkennen van gezichtsexpressies. De Psycholoog, 47(10), 10-17. Download.

Kret, M.E., & de Gelder, B. (2012). Islamic context influences how emotion is recognized from the eyes. Frontiers in Psychology, 3(110). I = 2.800. Download.

Kret, M.E., & de Gelder, B. (2012). A review on sex differences in processing emotions. Neuropsychologia, 50(7):1211-1221. I = 4.739. Download.


Kret, M.E. (2011). Context, gender and personality factors influencing the perception of facial and bodily expressions of emotion. Ridderkerk: Ridderprint. Download.

Kret, M.E., Grèzes, J., Pichon, S. & de Gelder, B. (2011). Men fear other men most: Gender specific brain activations in perceiving threat from dynamic faces and bodies. An fMRI study. Frontiers in Psychology, 2(3). I = 2.800. Download.

Kret, M.E. & de Gelder, B., Denollet, J. (2011). The role of negative affectivity and social inhibition in perceiving social threat: an fMRI study. Neuropsychologia, 49(5), 1187-1193. I = 4.739. Download.

Kret, M.E., Grèzes, J., Pichon, S. & de Gelder, B. (2011). Similarities and differences in perceiving threat from dynamic faces and bodies. An fMRI study. NeuroImage, 54(2), 1755- 1762. I = 6.817. Download.


Kret, M.E. & de Gelder, B. (2010). Social context influences recognition of bodily expressions. Experimental Brain Research, 203, 169-180. I = 2.296. Download.


de Gelder, B., Van den Stock, J., Meeren, H., Kret, M.E., Tamietto, M. (2009). Standing up for the body. Recent progress in uncovering the networks involved in processing bodies and bodily expressions. Neuroscience and Biobehavioral Reviews, 364, (1535), 3475-3484. I = 10.240. Download.


du Bois-Reymond, M., Leccardi, C., Magaraggia, S., Menz, S., Stauber, B., Biggart, A.,  Kovacheva, S., Manolova, P., Ule, M., Kuhar, M., Svab, A. & Kret, M.E. (2008). Young parenthood, agency and social change. Thematic report for the UP2YOUTH-project.


Kret, M.E. (in prep). Uitingen van emoties in mens en dier. Uitgeverij Atlas Contact, Amsterdam, the Netherlands.

Kret, M.E. (2011). Context, gender and personality factors, influencing the perception of facial and bodily expressions of emotion. Doctoral dissertation. Ridderkerk: Ridderprint. Download.

Book chapters

Kret, M.E., Sinke, C.B.A & de Gelder, B. (2011). Emotion perception and health. In Nyclicek, I., Vingerhoets, A.J.J.M. & Zeelenberg, M. (Eds.), Emotion regulation and well-being (pp.261-280). New York: Springer. Download.

Sinke, C.B.A., Kret, M.E. & de Gelder, B. (2010). Embodied perception of emotion. Psychology press. MINET: Measuring the impossible. Download.

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